Bergmann's rule is an ecogeographical principle stating that within broadly distributed taxonomic groups, larger species are found in colder environments and smaller species in warmer regions.Carl Bergmann, an anatomist and physiologist from Göttingen, published an article in 1847 titled 'On the relationship between heat economy in animals and their size.' He noticed that among closely related species of mammals and birds, those living in high latitudes are noticeably larger. Bergmann explained this with simple physics: heat production is proportional to body mass (volume), while heat loss is proportional to surface area. Since volume increases faster than area with increasing size, large animals have a more favorable ratio of heat production to heat loss. This allows them to survive in harsh climates. Later, ecologists confirmed the rule for many groups: from deer to penguins. Interestingly, Bergmann's rule often does not apply to insects and reptiles — they depend on external heat.
Carl Georg Lucas Christian Bergmann, professor of anatomy and physiology at the University of Göttingen, published the work 'Über die Verhältnisse der Wärmeökonomie der Thiere zu ihrer Grösse' in 1847. By comparing collections of stuffed animals from different regions, he was the first to quantitatively substantiate the link between climate and body size. Bergmann relied on Newton's laws of heat transfer and emerging ideas about metabolism in physiology. His conclusions were long disputed, but in the 20th century they were confirmed by mass measurements: from wolves in North America to wild boars in Eurasia. In 1937, David Lack showed that the rule also applies to birds when considering not only latitude but also altitude above sea level.
Bergmann's rule is used in paleoclimatology to estimate past temperatures from the sizes of fossil mammals. In agriculture, it is considered when acclimatizing livestock breeds: Holstein cows in Canada are larger than in Texas. The rule also helps predict the consequences of global warming: many species are expected to decrease in size. Limits of applicability: the rule does not apply to ectothermic animals (fish, reptiles), whose thermoregulation is behavioral rather than metabolic.
When breeding fur-bearing animals in northern latitudes, larger individuals are selected: their fur is denser and heat loss is lower, which increases the yield of high-quality raw material.
Emperor penguins, living in Antarctica, are the largest of all penguins: their height reaches 120 cm and weight up to 45 kg. Meanwhile, Galapagos penguins from the equator are half the size. Yet they manage to survive on scorching lava shores by hiding in the shade of rocks.





